いつもお世話になってます。久々にまた投稿させてください。
今日もSCにおける質問です。どなたか分かられることは解説お願いします。

2. In the most common procedure for harvesting forage crops such as alfalfa, as much as 20 percent of the leaf and small-stem material, <which is the most nutritious of all the parts of the plant, shattered and fell> to the ground.

(A) which is the most nutritious of all the parts of the plant, shattered and fell
(B) the most nutritious of all parts of the plant, shatter and fall
(C) the parts of the plant which were most nutritious, will shatter and fall
(D) the most nutritious parts of the plant, shatters and falls
(E) parts of the plant which are the most nutritious, have shattered and fallen

Ans.(D)
この場合において、"shatters and falls"の主語を何なのでしょうか？私は"as much as 20 percent of the leaf and small-stem material"を主語と認識しこれを複数形だと思い、正解である（D)を消したのですが。。。これは単数形なのでしょうか？

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